A visit to arguably the most famous archipelago on Earth, the Galapagos. It's home to a myriad of bizarre and unique creatures, endemic to these islands - but how did they get here and what is the key to these extraordinary islands that allows them to thrive? The programme reveals that this key holds not just the secret to life here, but also to how Darwin was able to leave with the ideas that would revolutionise biology.

The coast is the dynamic border between land and sea; powered by the tides and thrashed by waves, this is a world of continuous change. Opportunities never last long here, so hunters are always in a race against time. The coast is the only place on the planet where predators from air, land and sea come together. Dolphins that leave the safety of the sea to fish, walking octopuses, ingenious monkeys, fishing wolves and the greatest gathering of feeding humpback whales come to the coast to hunt. For all, timing is everything.

the final episode deals with plants that live in hostile environments. Attenborough visits Ellesmere Island, north of the Arctic Circle, to demonstrate that even in a place that is unconducive to life, it can be found. Algae and lichens grow in or on rock, and during summer, when the ice melts, flowers are much more apparent. However, they must remain close to the ground to stay out of the chilling wind. In the Tasmanian mountains, plants conserve heat by growing into 'cushions' that act as solar panels, with as many as a million individual shoots grouped together as one. Others, such as the lobelia in Mount Kenya, have a 'fur coat' of dense hairs on their leaves. The saguaro cactus in the Sonoran Desert flourishes because of its ability to retain vast amounts of water, which can't be lost through leaves because it has none. Many desert dwellers benefit from an accelerated life cycle, blooming rapidly within weeks after rainfall. Conversely, Mount Roraima is one of the wettest places on Earth. It is a huge sandstone plateau with high waterfalls and nutrients are continuously washed away, so plants have to adapt their diet if they are to survive. A bladderwort is shown invading a bromeliad. Inhabitants of lakes have other problems to contend with: those that dominate the surface will proliferate, and the Amazon water lily provides an apt illustration. Attenborough ends the series with an entreaty for the conservation of plant species.

The Milky Way, our galaxy, is a magnificent sight in the night sky, but we know surprisingly little about it for certain. What is its shape? How many stars does it actually contain? What lies at its centre? The Gaia space telescope will answer these questions, being armed with the most advanced camera to leave our planet, and it will allow us to see our galaxy as we've never seen it before. The Sky at Night visits the factory in Chelmsford that made the astonishing sensor at the heart of the mission.

Michael Ruppert is an independent journalist who has made a minor career out of telling people news that most folks do not want to know. Ruppert, a former police officer, predicted the Wall Street debacle of 2008 several years before the fact, at a time when most analysts were still imagining infinite growth for the stock market and major investment banks. Since then, his vision of the world's future has grown only darker. As Ruppert sees it, civilization and the global economy has yet to wean itself off fossil fuels, and when the world's supply of oil finally runs out, it will lead to a global financial catastrophe that will leave no one unscathed. But while most of what Ruppert has to say bears the ring of truth, there's a small audience for his dire message -- the primary medium for his work is a self-published newsletter, and his most recent book has done so poorly in the marketplace that he faces eviction from his home. Is Ruppert right? And if he is, why doesn't anyone care? Filmmaker Chris Smith profiles Michael Ruppert and gives him a chance to explain his apocalyptic vision of the future at length

The journeys of the Voyager probes is put in the context of the Netherlands in the seventeenth century, with a centuries-long tradition of sailing ship explorers, and its contemporary thinkers (such as Constantijn Huygens and his son Christian). Their discoveries are compared to the Voyager probes' discoveries among the Jovian and Saturn systems. In Cosmos Update, image processing reconstructs Voyager’s worlds and Voyager’s last portrait of the Solar System as it leaves is shown.